- large trees
- branches lacking spines
- pair of glands at top of petiole (base of blade)
- elongate raceme with 12-20 or more flowers
- leaves ovate to elliptic or lanceolate, widest at or below the middle, glossy above
- margins crenulate - the teeth incurved
- sepals glandless or with at most 2-4 glands
- rust-colored hairs along the midrib on the underside of the leaves